Reporter CRISPR screens decipher cis- and trans-regulatory principles at the Xist locus DOI Creative Commons
Till Schwämmle, Gemma Noviello,

Eleni Kanata

и другие.

bioRxiv (Cold Spring Harbor Laboratory), Год журнала: 2024, Номер unknown

Опубликована: Окт. 10, 2024

Abstract Developmental genes are controlled by an ensemble of cis -acting regulatory elements (REs), which in turn respond to multiple trans transcription factors (TFs). Understanding how a -regulatory landscape integrates information from many dynamically expressed TFs has remained challenge. We develop combined CRISPR-screening approach using endogenous RNA and RE-reporters as readouts. Applied the Xist locus, crucial for X-chromosome inactivation females, this method allows us comprehensively identify Xist-controlling map their TF-RE wiring. find group transiently that regulate proximal REs, driving binary activation expression. These basal activators more highly cells with two X chromosomes, potentially female-specific upregulation. A second set developmental is upregulated later during differentiation targets distal REs. This axis achieve high levels RNA, necessary inactivation. Our findings support model gene regulation targeting REs drive ON-OFF decisions, while interacting control output.

Язык: Английский

Chromatin and aberrant enhancer activity in KMT2A rearranged acute lymphoblastic leukemia DOI Creative Commons
Thomas A. Milne

Current Opinion in Genetics & Development, Год журнала: 2024, Номер 86, С. 102191 - 102191

Опубликована: Апрель 4, 2024

To make a multicellular organism, genes need to be transcribed at the right developmental stages and in tissues. DNA sequences termed 'enhancers' are crucial achieve this. Despite concerted efforts, exact mechanisms of enhancer activity remain elusive. Mixed lineage leukemia (MLL or KMT2A) rearrangements (MLLr), commonly observed cases acute lymphoblastic (ALL) myeloid leukemia, produce novel in-frame fusion proteins. Recent work has shown that MLL-AF4 protein drives aberrant key oncogenes ALL, dependent on continued presence complex components. As well as providing some general insights into function, these observations may also provide an explanation for transcriptional heterogeneity MLLr patients.

Язык: Английский

Процитировано

5

The α-globin super-enhancer acts in an orientation-dependent manner DOI Creative Commons
Mira Kassouf, Helena S. Francis, Matthew Gosden

и другие.

Nature Communications, Год журнала: 2025, Номер 16(1)

Опубликована: Янв. 25, 2025

Abstract Individual enhancers are defined as short genomic regulatory elements, bound by transcription factors, and able to activate cell-specific gene expression at a distance, in an orientation-independent manner. Within mammalian genomes, enhancer-like elements may be found individually or within clusters referred locus control regions super-enhancers (SEs). While these behave similarly individual with respect cell specificity, distribution their orientation-dependence has not been formally tested. Here, using the α-globin model, we show that while enhancer works manner, direction of activity SE changes its orientation. When is inverted normal chromosomal context, targets, genes, severely reduced normally silent genes lying upstream upregulated. These findings add our understanding enhancer-promoter specificity precisely transcription.

Язык: Английский

Процитировано

0

The roles of enhancer, especially super-enhancer-driven genes in tumor metabolism and immunity DOI
Shouwu Guo, Lu Zhang,

Jiao Ren

и другие.

International Journal of Biological Macromolecules, Год журнала: 2025, Номер unknown, С. 142414 - 142414

Опубликована: Март 1, 2025

Язык: Английский

Процитировано

0

Is Enhancer Function Driven by Protein–Protein Interactions? From Bacteria to Leukemia DOI Creative Commons
Nicholas T. Crump, Thomas A. Milne

BioEssays, Год журнала: 2025, Номер unknown

Опубликована: Апрель 8, 2025

ABSTRACT The precise regulation of the transcription genes is essential for normal development and maintenance life. Aberrant gene expression changes drive many human diseases. Despite this, we still do not completely understand how controlled in living systems. Enhancers are key regulatory elements that enable cells to specifically activate response environmental cues, or a stage tissue‐specific manner. Any model enhancer activity needs answer two main questions: (1) enhancers able identify act on specific (2) influence transcription. To address these points, first outline some basic principles can be established from simpler prokaryotic systems, then discuss recent work aberrant leukemia. We argue highly protein–protein interactions driver enhancer‐promoter proximity, allowing enhancer‐bound factors directly RNA polymerase

Язык: Английский

Процитировано

0

DNA G-quadruplex structures act as functional elements in α- and β-globin enhancers DOI Creative Commons

Colm Doyle,

Krzysztof Herka, Sean M. Flynn

и другие.

Genome biology, Год журнала: 2025, Номер 26(1)

Опубликована: Июнь 4, 2025

Abstract Background Enhancer elements interact with target genes at a distance to modulate their expression, but the molecular details of enhancer–promoter interaction are incompletely understood. G-quadruplex DNA secondary structures (G4s) have recently been shown co-occur 3D chromatin interactions; however, functional importance G4s within enhancers remains unclear. Results In this study, we identify novel G4 two locus control regions human α- and β-globin loci. We find that mutating motifs by genome editing prevents folding into in cells disrupts interactions gene expression manner comparable whole enhancer deletion. Furthermore, restoration structure formation using dissimilar G4-forming primary sequence recovers specific enhancer-gene expression. Through proteomic, biophysical, genomic profiling, tightly linked maintenance an active state RNA polymerase II recruitment regulate Conclusions Our study shows folded can act as mediate support enhancer-driven globin regulation.

Язык: Английский

Процитировано

0

A mammalian tripartite enhancer cluster controls hypothalamic Pomc expression, food intake, and body weight DOI
Daniela Rojo,

Clara E. Hael,

Agustina Soria

и другие.

Proceedings of the National Academy of Sciences, Год журнала: 2024, Номер 121(18)

Опубликована: Апрель 23, 2024

Food intake and energy balance are tightly regulated by a group of hypothalamic arcuate neurons expressing the proopiomelanocortin ( POMC) gene. In mammals, arcuate-specific POMC expression is driven two cis -acting transcriptional enhancers known as nPE1 nPE2. Because mutant mice lacking these still showed Pomc mRNA, we searched for additional elements contributing to expression. By combining molecular evolution with reporter gene in transgenic zebrafish mice, here, identified mammalian enhancer that named nPE3, carrying several binding sites also present nPE2 transcription factors activate neuronal expression, such ISL1, NKX2.1, ERα. We found nPE3 originated lineage leading placental mammals remained under purifying selection all orders, although it was lost Simiiformes (monkeys, apes, humans) following unique segmental deletion event. Interestingly, ablation from mouse genome led drastic reduction (>70%) mRNA during development only moderate (<33%) adult mice. Comparison between double (nPE1 nPE2) triple (nPE1, nPE2, nPE3) mutants revealed relative contribution its importance control food adiposity male female Altogether, results demonstrate integrates tripartite cluster partially redundant upon convergent evolutionary process critical body weight homeostasis.

Язык: Английский

Процитировано

0

Reporter CRISPR screens decipher cis- and trans-regulatory principles at the Xist locus DOI Creative Commons
Till Schwämmle, Gemma Noviello,

Eleni Kanata

и другие.

bioRxiv (Cold Spring Harbor Laboratory), Год журнала: 2024, Номер unknown

Опубликована: Окт. 10, 2024

Abstract Developmental genes are controlled by an ensemble of cis -acting regulatory elements (REs), which in turn respond to multiple trans transcription factors (TFs). Understanding how a -regulatory landscape integrates information from many dynamically expressed TFs has remained challenge. We develop combined CRISPR-screening approach using endogenous RNA and RE-reporters as readouts. Applied the Xist locus, crucial for X-chromosome inactivation females, this method allows us comprehensively identify Xist-controlling map their TF-RE wiring. find group transiently that regulate proximal REs, driving binary activation expression. These basal activators more highly cells with two X chromosomes, potentially female-specific upregulation. A second set developmental is upregulated later during differentiation targets distal REs. This axis achieve high levels RNA, necessary inactivation. Our findings support model gene regulation targeting REs drive ON-OFF decisions, while interacting control output.

Язык: Английский

Процитировано

0