Nonequilibrium phases of a biomolecular condensate facilitated by enzyme activity DOI Creative Commons
Sebastian Coupe, Nikta Fakhri

bioRxiv (Cold Spring Harbor Laboratory), Год журнала: 2024, Номер unknown

Опубликована: Авг. 11, 2024

Abstract Biomolecular condensates represent a frontier in cellular organization, existing as dynamic materials driven out of equilibrium by active processes. Here we explore mechanisms condensate regulation examining the interplay between DEAD-box helicase activity and RNA base-pairing interactions within ribonucleoprotein condensates. We demonstrate how ATP-dependent helicases—a key class enzymes regulation—acts nonequilibrium driver properties through continuous remodeling interactions. By combining LAF-1 with designer hairpin concatemer, unveil complex landscape behaviors, including time-dependent alterations partitioning, evolving morphologies, shifting dynamics. Importantly, reveal an antagonistic relationship secondary structure which promotes homogeneity via steady state. elucidating these mechanisms, gain deeper understanding organization expand potential for synthetic systems.

Язык: Английский

Reaction-Driven Diffusiophoresis of Liquid Condensates: Potential Mechanisms for Intracellular Organization DOI Creative Commons
G. S. Hafner, Marcus Müller

ACS Nano, Год журнала: 2024, Номер 18(26), С. 16530 - 16544

Опубликована: Июнь 14, 2024

The cellular environment, characterized by its intricate composition and spatial organization, hosts a variety of organelles, ranging from membrane-bound ones to membraneless structures that are formed through liquid–liquid phase separation. Cells show precise control over the position such condensates. We demonstrate organelle movement in external concentration gradients, diffusiophoresis, is distinct one colloids because fluxes can remain finite inside liquid-phase droplets latter arises incompressibility. Within domains diffusiophoresis naturally biochemical reactions driven chemical fuel produce waste. Simulations analytical arguments within minimal model reaction-driven separation reveal directed stems two contributions: Fuel waste refilled or extracted at boundary, resulting which (i) induce product via incompressibility (ii) result an asymmetric forward reaction droplet's surroundings (as well as backward droplet), thereby shifting position. former contribution dominates sets direction movement, toward away source sink, depending on molecules' affinity waste, respectively. mechanism thus provides simple means organize condensates with different composition. Particle-based simulations systems more complex cycles corroborate robustness universality this mechanism.

Язык: Английский

Процитировано

5

Charge separation at liquid interfaces DOI Creative Commons
Arghya Majee, Christoph A. Weber, Frank Jülicher

и другие.

Physical Review Research, Год журнала: 2024, Номер 6(3)

Опубликована: Авг. 5, 2024

We present a theory for phase-separated liquid coacervates with salt, taking into account spatial heterogeneities and interfacial profiles. find that charged layers of alternating sign can form around the interface while bulk phases remain approximately charge neutral. show salt concentration regulates number amplitude layer's density electrostatic potential. Such either repel or attract single-charged molecules diffusing across interface. Our could be relevant artificial systems biomolecular condensates in cells. work suggests interfaces mediate charge-specific transport similar to membrane-bound compartments. Published by American Physical Society 2024

Язык: Английский

Процитировано

5

Bursting of condensates DOI Creative Commons
Isabell Tunn, Grégory Beaune, Jennifer Tersteegen

и другие.

Communications Physics, Год журнала: 2024, Номер 7(1)

Опубликована: Май 17, 2024

Abstract Numerous biomolecular shell-forming condensates are reported in cells and bioengineered vitro. The relationship between the molecular structure of their biophysical properties remains largely unexplored. To fill this gap, we characterize spider silk proteins based on Araneus diadematus major ampulla gland fibroin 3 (ADF3) using micropipette aspiration. We observe that can burst during aspiration like soap bubbles or polymersomes, demonstrating formation a dense protein layer (shell) at condensate interface. tendency to is more pronounced for formed from with weakly dimerizing terminal blocks. develop model analyse bursting condensates, obtain surface bulk viscosity, estimate shell thickness viscosity. Understanding controlling will open avenues use materials, as compartments reactions drug delivery systems.

Язык: Английский

Процитировано

4

Reactive Brownian Dynamics of Chemically Fueled Droplets: Roles of Attraction and Deactivation Modes DOI
Lennard Holschuh, Joachim Dzubiella

The Journal of Physical Chemistry B, Год журнала: 2025, Номер 129(2), С. 626 - 636

Опубликована: Янв. 6, 2025

The self-assembly of biological membraneless organelles can be mimicked by active droplets resulting from chemically fueled microphase separation. However, how the nonequilibrium, transient structure these controlled through physicochemical input parameters is not yet well understood. In our work, a two-state chemical reaction and subsequent droplet growth decay are modeled with reactive Brownian dynamics simulation in two spatial dimensions. model, particles that activated via consumption fuel become attractive accumulate into droplets. A local-density-dependent distinction droplet's 'internal' 'external' allows for structural feedback giving further control over deactivation process. shows only external slows down stabilizes droplets, whereas internal lead to temporary encapsulation deactivated (in nonequilibrium 'core–shell' structures) where serve as an outer shell. Additionally, role hydrophobicity resembled attraction energy ε dependency formation on various investigated. For example, high finite-size crystalline while other parameter choices indicate bimodal size distributions at specific times. Similarities differences related experiments discussed.

Язык: Английский

Процитировано

0

Complex dynamics in reaction-phase separation systems DOI
Dino Osmanović, Elisa Franco

Physical review. E, Год журнала: 2025, Номер 111(2)

Опубликована: Фев. 7, 2025

We investigate the emergence of sustained spatiotemporal behaviors in reaction-phase separation systems. focus on binary systems, which either one or both species can phase separate, and we discuss stability homogeneous state determining conditions for a Hopf-type bifurcation. then examine effects specific autocatalytic chemical reaction, computationally determine full solutions to partial differential equations. find that when pulsed dynamics arise dimension. When considered two dimensions, system generates persistent, complex dynamic droplets, do not generally appear if only separate. finally with features, be understood using framework cellular automata.

Язык: Английский

Процитировано

0

Chemically active wetting DOI Creative Commons
Susanne Liese, Xueping Zhao, Christoph A. Weber

и другие.

Proceedings of the National Academy of Sciences, Год журнала: 2025, Номер 122(15)

Опубликована: Апрель 9, 2025

Wetting of liquid droplets on passive surfaces is ubiquitous in our daily lives, and the governing physical laws are well understood. When become active, however, wetting remain elusive. Here, we propose chemically active as a class systems where surface due to binding process that maintained away from equilibrium. We derive corresponding nonequilibrium thermodynamic theory show fundamentally changes behavior, leading steady, states with droplet shapes reminiscent pancake or mushroom. The origin such anomalous can be explained by mapping electrostatics, pairs sinks sources correspond electrostatic dipoles along triple line. This an example more general analogy, localized chemical activity gives rise multipole field potential. underlying physics relevant for cells, droplet-forming proteins bind membranes accompanied turnover biological fuels.

Язык: Английский

Процитировано

0

Active RNA synthesis patterns nuclear condensates DOI Creative Commons
Salman F. Banani, Andriy Goychuk, Pradeep Natarajan

и другие.

bioRxiv (Cold Spring Harbor Laboratory), Год журнала: 2024, Номер unknown

Опубликована: Окт. 13, 2024

Biomolecular condensates are membraneless compartments that organize biochemical processes in cells. In contrast to well-understood mechanisms describing how form and dissolve, the principles underlying condensate patterning - including their size, number spacing cell remain largely unknown. We hypothesized RNA, a key regulator of formation dissolution, influences patterning. Using nucleolar fibrillar centers (FCs) as model condensate, we found inhibiting ribosomal RNA synthesis significantly alters FCs. Physical theory experimental observations support whereby active generates non-equilibrium state arrests coarsening thus contributes Altering FC by expression component TCOF1 impairs processing, linking biological function. These results reveal states driven chemical regulate patterning, which is important for cellular biochemistry

Язык: Английский

Процитировано

3

Anomalous Fluctuations in a Droplet of Chemically Active Colloids or Enzymes DOI Creative Commons
K. R. Prathyusha, Suropriya Saha, Ramin Golestanian

и другие.

Physical Review Letters, Год журнала: 2024, Номер 133(5)

Опубликована: Июль 30, 2024

Chemically active colloids or enzymes cluster into dense droplets driven by their phoretic response to collectively generated chemical gradients. Employing Brownian dynamics simulation techniques, our study of the such a chemically droplet uncovers rich variety structures and dynamical properties, including full range fluidlike solidlike behavior, non-Gaussian positional fluctuations. Our work sheds light on complex constituents metabolic clusters, which are main drivers nonequilibrium activity in living systems.

Язык: Английский

Процитировано

1

Polarised crowd in motion: Insights into statistical and dynamical behavior DOI Creative Commons

Pratikshya Jena,

Shradha Mishra

Research Square (Research Square), Год журнала: 2024, Номер unknown

Опубликована: Апрель 26, 2024

Abstract The collection of active agents often exhibits intriguing statistical and dynamical properties, particularly when consid- ering human crowds. In this study, we have developed a computational model to simulate the recent experiment on real marathon races by Bain et al., [Science 363, 46–49 (2019)]. Our primary goal is investigate impact race staff crowd dynamics. By comparing simulated with without presence staff, our study reveals that local velocity density participants display wave pattern akin for both cases. observed traveling in consistently propagates at constant speed, regardless system size under eration. participants’ dynamics longitudinal direction primarily contribute fluctuations, while fluctuations transverse are suppressed. absence weaken significantly affecting other dynamic characteristics crowd. Through research, aim deepen understanding motion, providing insights can inform development effective management strategies successful control such events.

Язык: Английский

Процитировано

0

Nonequilibrium phases of a biomolecular condensate facilitated by enzyme activity DOI Creative Commons
Sebastian Coupe, Nikta Fakhri

bioRxiv (Cold Spring Harbor Laboratory), Год журнала: 2024, Номер unknown

Опубликована: Авг. 11, 2024

Abstract Biomolecular condensates represent a frontier in cellular organization, existing as dynamic materials driven out of equilibrium by active processes. Here we explore mechanisms condensate regulation examining the interplay between DEAD-box helicase activity and RNA base-pairing interactions within ribonucleoprotein condensates. We demonstrate how ATP-dependent helicases—a key class enzymes regulation—acts nonequilibrium driver properties through continuous remodeling interactions. By combining LAF-1 with designer hairpin concatemer, unveil complex landscape behaviors, including time-dependent alterations partitioning, evolving morphologies, shifting dynamics. Importantly, reveal an antagonistic relationship secondary structure which promotes homogeneity via steady state. elucidating these mechanisms, gain deeper understanding organization expand potential for synthetic systems.

Язык: Английский

Процитировано

0