Cladistics,
Journal Year:
2021,
Volume and Issue:
37(5), P. 630 - 638
Published: May 28, 2021
Abstract
We
recently
published
a
method
to
infer
ancestral
landmark‐based
shape
ontogenies
that
takes
into
account
the
possible
existence
of
changes
in
developmental
timing.
Here
we
describe
SPASOS,
software
perform
analysis.
SPASOS
is
an
open‐source
Windows
program
written
C.
Input
data
include
landmark
coordinates
for
each
specimen
–with
corresponding
information
about
timing–
and
phylogenetic
tree
showing
relationships
among
species
sampled.
As
output,
produces
image
files
easy
visualization
results
useful
post‐processing.
The
incorporates
interpolating
function,
based
on
weighting
moving
averages,
which
allows
analysis
with
scarce
along
ontogenetic
trajectory.
An
empirical
evaluation
this
function
showed
its
suitability
fill
incomplete
trajectories.
Finally,
present
reanalysis
dataset,
where
timing
were
originally
inferred
by
considering
PCA
scores
as
variables.
Both
approaches
retrieved
same
four
largest
timing,
but
differed
shapes
inferred.
Here
we
report
a
new
articulated
skeleton
of
Yamaceratops
dorngobiensis
(MPC-D
100/553)
from
the
Khugenetjavkhlant
locality
at
Shine
Us
Khudag
(Javkhlant
Formation,
?Santonian-Campanian)
eastern
Gobi
Desert,
Mongolia,
which
represents
first
substantially
complete
and
juvenile
individual
this
taxon.The
specimen
includes
nearly
cranium
large
portions
vertebral
column
appendicular
skeleton.Its
skull
is
about
2/3
size
holotype
specimen,
based
on
mandibular
length.Its
ontogenetic
stage
confirmed
by
multiple
indicators
skeletal
morphological
immaturity
known
in
ceratopsians,
such
as
long-grained
surface
texture
long
bones,
smooth
external
postorbital,
open
neurocentral
sutures
all
caudal
vertebrae,
orbit
relative
to
postorbital
jugal,
low
angle
lacrimal
ventral
ramus
maxillary
teeth
row,
narrow
frontal,
straight
edge
dentary.Osteohistological
analysis
MPC-D
100/553
recovered
three
lines
arrested
growth,
implying
around
years
age
when
it
died,
verified
specimen's
immature
stage.The
adds
autapomorphy
Yamaceratops,
anteroventral
margin
fungiform
dorsal
end
being
excluded
antorbital
fossa.Furthermore,
shows
unique
combination
diagnostic
features
some
other
basal
neoceratopsians:
ventrally
hooked
rostral
bone
Aquilops
americanus
very
tall
middle
neural
spines
or
more
than
four
times
high
centrum
Koreaceratops
hwaseongensis,
Montanoceratops
cerorhynchus,
Protoceratops
andrewsi.The
jugal
with
subtemporal
Here
we
report
a
new
articulated
skeleton
of
Yamaceratops
dorngobiensis
(MPC-D
100/553)
from
the
Khugenetjavkhlant
locality
at
Shine
Us
Khudag
(Javkhlant
Formation,
?Santonian-Campanian)
eastern
Gobi
Desert,
Mongolia,
which
represents
first
substantially
complete
and
juvenile
individual
this
taxon.The
specimen
includes
nearly
cranium
large
portions
vertebral
column
appendicular
skeleton.Its
skull
is
about
2/3
size
holotype
specimen,
based
on
mandibular
length.Its
ontogenetic
stage
confirmed
by
multiple
indicators
skeletal
morphological
immaturity
known
in
ceratopsians,
such
as
long-grained
surface
texture
long
bones,
smooth
external
postorbital,
open
neurocentral
sutures
all
caudal
vertebrae,
orbit
relative
to
postorbital
jugal,
low
angle
lacrimal
ventral
ramus
maxillary
teeth
row,
narrow
frontal,
straight
edge
dentary.Osteohistological
analysis
MPC-D
100/553
recovered
three
lines
arrested
growth,
implying
around
years
age
when
it
died,
verified
specimen's
immature
stage.The
adds
autapomorphy
Yamaceratops,
anteroventral
margin
fungiform
dorsal
end
being
excluded
antorbital
fossa.Furthermore,
shows
unique
combination
diagnostic
features
some
other
basal
neoceratopsians:
ventrally
hooked
rostral
bone
Aquilops
americanus
very
tall
middle
neural
spines
or
more
than
four
times
high
centrum
Koreaceratops
hwaseongensis,
Montanoceratops
cerorhynchus,
Protoceratops
andrewsi.The
jugal
with
subtemporal
Cladistics,
Journal Year:
2021,
Volume and Issue:
37(5), P. 630 - 638
Published: May 28, 2021
Abstract
We
recently
published
a
method
to
infer
ancestral
landmark‐based
shape
ontogenies
that
takes
into
account
the
possible
existence
of
changes
in
developmental
timing.
Here
we
describe
SPASOS,
software
perform
analysis.
SPASOS
is
an
open‐source
Windows
program
written
C.
Input
data
include
landmark
coordinates
for
each
specimen
–with
corresponding
information
about
timing–
and
phylogenetic
tree
showing
relationships
among
species
sampled.
As
output,
produces
image
files
easy
visualization
results
useful
post‐processing.
The
incorporates
interpolating
function,
based
on
weighting
moving
averages,
which
allows
analysis
with
scarce
along
ontogenetic
trajectory.
An
empirical
evaluation
this
function
showed
its
suitability
fill
incomplete
trajectories.
Finally,
present
reanalysis
dataset,
where
timing
were
originally
inferred
by
considering
PCA
scores
as
variables.
Both
approaches
retrieved
same
four
largest
timing,
but
differed
shapes
inferred.