Ornithischians
form
a
large
clade
of
globally
distributed
Mesozoic
dinosaurs,
and
one
their
three
major
radiations.Throughout
evolutionary
history,
exceeding
134
million
years,
ornithischians
evolved
considerable
morphological
disparity,
expressed
especially
through
the
cranial
osteodermal
features
most
distinguishable
representatives.The
nearly
two-century-long
research
history
on
has
resulted
in
recognition
numerous
diverse
lineages,
many
which
have
been
named.Following
formative
publications
establishing
theoretical
foundation
phylogenetic
nomenclature
throughout
1980s
1990s,
proposed
names
ornithischian
clades
were
provided
with
definitions.Some
these
definitions
proven
useful
not
changed,
beyond
way
they
formulated,
since
introduction.Some
names,
however,
multiple
definitions,
making
application
ambiguous.Recent
implementation
International
Code
Phylogenetic
Nomenclature
(ICPN,
or
PhyloCode)
offers
opportunity
to
explore
utility
previously
established
taxon
names.Since
Articles
ICPN
are
be
applied
retroactively,
all
published
prior
its
remain
informal
(and
ineffective)
light
Code.Here,
we
revise
dinosaur
clades;
revisit
76
preexisting
review
recent
historical
use,
formally
establish
definitions.Additionally,
introduce
five
new
names;
two
for
robustly
supported
later-diverging
hadrosaurids
ceratopsians,
uniting
Paleontological Research,
Journal Year:
2024,
Volume and Issue:
28(4)
Published: July 23, 2024
Of
the
four
main
Japanese
islands,
Late
Cretaceous
dinosaur
fossils
have
been
discovered
in
Hokkaido,
Honshu,
and
Kyushu.
Here,
we
report
first
from
remaining
island
Shikoku,
an
isolated
dorsal
vertebra
upper
Campanian
Hiketa
Formation
(Izumi
Group)
Sanuki,
Kagawa
Prefecture.
The
is
incomplete,
comprising
a
robust
centrum
with
subrectangular
profile
that
generally
characteristic
of
dinosaurs.
Histological
examination
reveals
moderately
packed
trabecular
bone
complete
absence
pneumatic
structures,
which
precludes
referral
specimen
to
Saurischia
or
Pterosauria.
Instead,
interpret
it
as
hadrosauriform
ornithischian
based
on
slightly
expanded
anterior
articular
surface
centrum;
heart-shaped
outline
intervertebral
surface,
resulting
its
dorsally
wide
ventrally
narrow
shape;
ventral
keel;
neural
canal
partially
embedded
into
centrum.
As
positioning
common
Bactrosaurus
higher
taxa
within
Hadrosauriformes,
tentatively
refer
Hadrosauroidea.
This
discovery
provides
additional
evidence
hadrosauriforms
had
dispersed
present-day
Japan
by
highlights
potential
provide
improve
our
understanding
diversity
at
extreme
eastern
continental
margin
Eurasia.
Зоологический журнал,
Journal Year:
2024,
Volume and Issue:
103(6), P. 15 - 30
Published: Dec. 3, 2024
The
determination
of
individual
age
and
ontogenetic
stage
(juvenile,
subadult,
adult)
fossil
vertebrates
is
important
for
the
initial
taxonomic
affiliation,
as
well
further
evolutionary
paleobiological
interpretations.
Determination
(=
relative
age)
carried
out
by
various
methods,
including
paleohistological
analysis.
study
thin
sections
tetrapod
bones
allows
us
to
assume
how
many
years
animal
lived
(skeletochronological
method)
determine
according
a
set
age-related
histological
markers:
change
in
type
bone
matrix
vascularization,
distance
between
growth
marks,
formation
external
fundamental
system
(EFS),
inner
(endosteal)
outer
(periosteal)
circumferential
layers
(OCL,
ICL),
occurrences
secondary
remodeling
–
Haversian
substitution
trabeculae.
Depending
on
phylogenetic
position
biological
peculiarities
group
tetrapods,
“histologic
markers”
may
be
different.
Journal of Vertebrate Paleontology,
Journal Year:
2022,
Volume and Issue:
42(5)
Published: Nov. 25, 2022
A
metatarsal
IV
from
a
hitherto
unknown
endemic
small-bodied
styracosternan
is
described
uppermost
Maastrichtian
strata
of
NE
Spain,
part
the
Ibero-Armorican
island
Late
Cretaceous
European
Archipelago.
This
element
unique
among
fourth
metatarsals
all
other
ornithopods
in
combining
greatly
elongated
proportions
basally
branching
members
clade
with
prominent
medial
flange
seen
styracosternans.
specimen
becomes
holotype
new
genus
and
species.
Histological
data
indicate
that
this
individual
was
late
subadult
at
time
death,
but
not
far
reaching
somatic
maturity.
By
analogy
elongate
lightly
built,
cursorial
ornithopods,
it
likely
animal
also
capable
rapid
locomotion.
consistent
differential
distribution
intense
bone
remodeling
IV,
resulting
biomechanical
stress
produced
during
hindlimb
propelling.
Notably,
represents
an
exception
to
mediportality
large
body
size
characterize
clade.
The
elongation
inferred
cursoriality,
small
constitute
case
evolutionary
convergence
distantly
related
non-iguanodontian
dryosaurid
elasmarian
iguanodontians.
Unlike
regions
world
where,
latest
stages
Cretaceous,
herbivore
ecological
niche
represented
by
non-iguanodontians
non-hadrosauriforms,
occupied
styracosternan.
This
project
seeks
to
investigate
the
morphology,
ontogeny,
and
evolution
of
skull
roof
braincase
characters
in
hadrosaurids
from
Campanian
northern
Laramidia
(Alberta
Montana),
a
setting
which
they
have
been
historically
recognized
as
having
high
abundance
diversity.
New
material
is
evaluated
clarify
distribution
informativeness
morphological
character
states,
allow
further
testing
previous
hypotheses
about
how
ontogenetic
development
these
evolved
within
clades,
enable
identification
previously
unrecorded
taxa.
Five
partial
skulls
Maiasaura
peeblesorum
are
described
bone
bed
Two
Medicine
Formation
Montana,
providing
basis
for
description
morphology
this
taxon,
an
series
allowing
crest
be
compared
related
new
middle
unit
Oldman
at
Milk
River
Ridge
Reservoir
near
Warner,
southern
Alberta
both
represent
first
occurrences
taxon
that
unit.
The
diagnostic
occurrence
Canada.
Brachylophosaurus
example
co-occurring
brachylophosaurins.
second
Parasaurolophini,
oldest
lambeosaurine
Alberta.
A
phylogenetic
analysis
using
maximum
parsimony
Bayesian
methods
supports
identifications,
also
re-identifies
specimen
Dinosaur
Park
geologically
youngest
brachylophosaurin
Hadrosaurid
(duck-billed)
dinosaur
bonebeds
are
exceedingly
prevalent
in
upper
Cretaceous
(Campanian-Maastrichtian)
strata
from
the
Midwest
of
North
America
(especially
Alberta,
Canada,
and
Montana,
U.S.A)
but
less
frequently
documented
more
northern
regions.The
Wapiti
Formation
northwestern
Alberta
is
a
largely
untapped
resource
terrestrial
palaeontological
information
missing
southern
due
to
deposition
marine
Bearpaw
Formation.In
2018,
Boreal
Dinosaur
Project
rediscovered
Spring
Creek
Bonebed,
which
had
been
lost
since
2002,
along
bank
River,
southwest
Grande
Prairie.Earlier
excavations
observations
Bonebed
suggested
that
site
yielded
young
hadrosaurines.Continued
work
2018
2019
recovered
~300
specimens
included
minimum
eight
individuals,
based
on
number
right
humeri.The
morphology
several
cranial
elements
unequivocally
supports
lambeosaurine
affinities,
making
sample
first
occurrence
lambeosaurines
Formation.The
overall
size
range
histology
bones
found
at
indicate
these
animals
were
uniformly
late
juveniles,
suggesting
age
segregation
was
life
history
strategy
among
hadrosaurids.Given
considerable
attained
by
lambeosaurines,
they
probably
segregated
breeding
population
during
nesting
or
caring
for
young,
rather
than
different
diet
locomotory
requirements.Dynamic
aspects
history,
such
as
segregation,
may
well
have
contributed
highly
diverse
cosmopolitan
nature
Late
hadrosaurids.
Hadrosaurid
(duck-billed)
dinosaur
bonebeds
are
exceedingly
prevalent
in
upper
Cretaceous
(Campanian-Maastrichtian)
strata
from
the
Midwest
of
North
America
(especially
Alberta,
Canada,
and
Montana,
U.S.A)
but
less
frequently
documented
more
northern
regions.The
Wapiti
Formation
northwestern
Alberta
is
a
largely
untapped
resource
terrestrial
palaeontological
information
missing
southern
due
to
deposition
marine
Bearpaw
Formation.In
2018,
Boreal
Dinosaur
Project
rediscovered
Spring
Creek
Bonebed,
which
had
been
lost
since
2002,
along
bank
River,
southwest
Grande
Prairie.Earlier
excavations
observations
Bonebed
suggested
that
site
yielded
young
hadrosaurines.Continued
work
2018
2019
recovered
~300
specimens
included
minimum
eight
individuals,
based
on
number
right
humeri.The
morphology
several
cranial
elements
unequivocally
supports
lambeosaurine
affinities,
making
sample
first
occurrence
lambeosaurines
Formation.The
overall
size
range
histology
bones
found
at
indicate
these
animals
were
uniformly
late
juveniles,
suggesting
age
segregation
was
life
history
strategy
among
hadrosaurids.Given
considerable
attained
by
lambeosaurines,
they
probably
segregated
breeding
population
during
nesting
or
caring
for
young,
rather
than
different
diet
locomotory
requirements.Dynamic
aspects
history,
such
as
segregation,
may
well
have
contributed
highly
diverse
cosmopolitan
nature
Late
hadrosaurids.
Hadrosaurid
(duck-billed)
dinosaur
bonebeds
are
exceedingly
prevalent
in
upper
Cretaceous
(Campanian-Maastrichtian)
strata
from
the
Midwest
of
North
America
(especially
Alberta,
Canada,
and
Montana,
U.S.A)
but
less
frequently
documented
more
northern
regions.The
Wapiti
Formation
northwestern
Alberta
is
a
largely
untapped
resource
terrestrial
palaeontological
information
missing
southern
due
to
deposition
marine
Bearpaw
Formation.In
2018,
Boreal
Dinosaur
Project
rediscovered
Spring
Creek
Bonebed,
which
had
been
lost
since
2002,
along
bank
River,
southwest
Grande
Prairie.Earlier
excavations
observations
Bonebed
suggested
that
site
yielded
young
hadrosaurines.Continued
work
2018
2019
recovered
~300
specimens
included
minimum
eight
individuals,
based
on
number
right
humeri.The
morphology
several
cranial
elements
unequivocally
supports
lambeosaurine
affinities,
making
sample
first
occurrence
lambeosaurines
Formation.The
overall
size
range
histology
bones
found
at
indicate
these
animals
were
uniformly
late
juveniles,
suggesting
age
segregation
was
life
history
strategy
among
hadrosaurids.Given
considerable
attained
by
lambeosaurines,
they
probably
segregated
breeding
population
during
nesting
or
caring
for
young,
rather
than
different
diet
locomotory
requirements.Dynamic
aspects
history,
such
as
segregation,
may
well
have
contributed
highly
diverse
cosmopolitan
nature
Late
hadrosaurids.