Weak link or strong foundation? Vulnerability of fine root networks and stems to xylem embolism DOI Creative Commons
Beatrice L. Harrison Day, Craig R. Brodersen, Timothy J. Brodribb

et al.

New Phytologist, Journal Year: 2024, Volume and Issue: unknown

Published: Sept. 12, 2024

Resolving the position of roots in whole-plant hierarchy drought-induced xylem embolism resistance is fundamental for predicting when species become isolated from soil water resources. Published research generally suggests that are most vulnerable organ plant vascular system, although estimates vary significantly. However, our knowledge root excludes fine (< 2 mm diameter) form bulk total absorptive surface area network and nutrient uptake. We measured stem vulnerability 10 major land clades (five angiosperms, three conifers, a fern lycophyte), using standardised situ methods (Optical Methods MicroCT). Mean across matched or exceeded stems all study species. In six these (one fern, one lycophyte, conifers angiosperm), were significantly more resistant than stems. No clear relationship was found between conduit diameter vulnerability. These results provide insight into hydraulic pathway at site uptake, challenge long-standing assumption

Language: Английский

Stomatal response to VPD is not triggered by changes in soil–leaf hydraulic conductance in Arabidopsis or Callitris DOI Creative Commons
Ibrahim Bourbia, Timothy J. Brodribb

New Phytologist, Journal Year: 2024, Volume and Issue: 242(2), P. 444 - 452

Published: Feb. 23, 2024

Stomatal closure under high VPD

Language: Английский

Citations

9

Variation pattern in the macromolecular (cellulose, hemicelluloses, lignin) composition of cell walls in Pinus tabulaeformis tree trunks at different ages as revealed using multiple techniques DOI
Weiwei Shen, Chen Zhang, Guangchao Wang

et al.

International Journal of Biological Macromolecules, Journal Year: 2024, Volume and Issue: 268, P. 131619 - 131619

Published: May 1, 2024

Language: Английский

Citations

7

Long‐Term in vivo Observation of Maize Leaf Xylem Embolism, Transpiration and Photosynthesis During Drought and Recovery DOI Creative Commons

Brendan S. Allen,

Jared J. Stewart, Stephanie K. Polutchko

et al.

Plant Cell & Environment, Journal Year: 2025, Volume and Issue: unknown

Published: Feb. 3, 2025

ABSTRACT Plant water transport is essential to maintain turgor, photosynthesis and growth. Water transported in a metastable state under large negative pressures, which can result embolism, that is, the loss of function by replacement liquid xylem sap with gas, as consequence stress. To avoid experimental artefacts, we used an optical vulnerability system quantify embolism occurrence across six fully expanded maize leaves characterize sequence physiological responses (photosynthesis, chlorophyll fluorescence, whole‐plant transpiration leaf inter‐vein distance) relation declining availability during severe Additionally, recovery presence sustained 6‐day period. Embolism formation occurred after other processes were substantially depressed irreversible upon rewatering. Recovery transpiration, net CO 2 assimilation photosystem II efficiency aligned severity whereas these traits returned near pre‐stress levels absence embolism. A better understanding relationships between downstream stress critical for improvement crop productivity resilience.

Language: Английский

Citations

0

Seasonal Shifts in Tree Water Use and Non‐Structural Carbohydrate Storage in a Tropical Dry Forest DOI Open Access
Maria Medeiros, André Luiz Alves de Lima, José Raliuson Inácio Silva

et al.

Plant Cell & Environment, Journal Year: 2025, Volume and Issue: unknown

Published: March 2, 2025

ABSTRACT Predictions of increased drought frequency and intensity have the potential to threaten forest globally. The key trees response is an understanding tree water use carbohydrate storage. Our objective was evaluate sap velocity dynamics non‐structural carbohydrates (NSC) in native a dry tropical forest, during rainy periods. We evaluated six species Caatinga: three deciduous with low wood density (WD), two high WD one evergreen measured velocity, xylem potential, stomatal conductance, phenology NSC. found that specie had higher frequent NSC production. While showed store mainly stem roots, leaf sprouting flowering at end period. also however, stored These results suggest under longer seasons irregular seasons, part resprout still season may be most affected.

Language: Английский

Citations

0

Stress dose explains drought recovery in Norway spruce DOI Creative Commons
Timo Knüver, Andreas Bär, Elias Hamann

et al.

Frontiers in Plant Science, Journal Year: 2025, Volume and Issue: 16

Published: March 6, 2025

Understanding the stress recovery of trees, particularly with respect to increasing droughts due climate change, is crucial. An often-overlooked aspect how short versus long drought events high intensity (i.e., low and dose) result in damage affect post-stress recovery. This study examines dynamics 3-year-old Picea abies following a (n = 5) 18 days or 9) 51 during late summer. We assessed canopy conductance tree transpiration linked i) terms minimum water potential, ii) duration inferred by below potential related 12% hydraulic loss (dP12), iii) dose cumulative deficit on P12 (TWDP12) as well (Ψcum), iv) percent conductive xylem area (PLA). Both treatments resulted stem root embolism higher PLA 49% ± 10% treatment compared 18% 6% consistent across measured plant parts. Suffering from leaf shedding (long drought, 32%; 12%), recovered 41% 3% control 66% 4% at 12 after release. These rates were explained observed (R2 0.66) dP12 0.62) but best metrics, TWDP12 0.88). Our highlights that should be integrated assess rates. Here, derived point dendrometers appears promising, it provides non-destructive temporal resolution dose.

Language: Английский

Citations

0

Sap flow through partially embolized xylem vessel networks DOI
Anna L. Jacobsen, Martín Venturas, Uwe G. Hacke

et al.

Plant Cell & Environment, Journal Year: 2024, Volume and Issue: 47(9), P. 3375 - 3392

Published: June 3, 2024

Abstract Sap is transported through numerous conduits in the xylem of woody plants along path from soil to leaves. When all are functional, vessel lumen diameter a strong predictor hydraulic conductivity. As vessels become embolized, sap movement becomes increasingly affected by factors operating at scales beyond individual conduits, creating resistances that result conductivity diverging diameter‐based estimates. These effects include pit resistances, connectivity, length, network topology, and or sector isolation. The impact these varies with level distribution emboli within network, manifest as alterations relationship between number embolized measured declines across vulnerability embolism curves. Divergences estimates reveal functional differences arise because species‐ tissue‐specific structures. Such divergences not uniform, tissues may diverge different ways differing degrees. Plants regularly operate under nonoptimal conditions contain conduits. Understanding implications function partially networks critical gaps our understanding occurring natural environments.

Language: Английский

Citations

2

Enhanced drought resistance in tomato via reduced auxin sensitivity: delayed dehydration and improved leaf resistance to embolism DOI Creative Commons
Moab T. Andrade, Amanda Á. Cardoso, Leonardo A. Oliveira

et al.

Physiologia Plantarum, Journal Year: 2024, Volume and Issue: 176(3)

Published: May 1, 2024

Abstract Auxins are master regulators of plant development and auxin perception mutants display smaller leaves, lower transpiration, narrower xylem vessels than their corresponding wild types. Here, we evaluated whether the leaf embolism resistance overall to drought altered in mutant diageotropica ( dgt ). Our assessments demonstrate that tomato exhibit considerably stems (‐24%), petioles (‐43%), midribs (‐34%) type. Alongside vessels, exhibited greater cell wall thickness‐to‐conduit diameter The water potential at 50% cumulative (P 50 ) type was ‐1.39 ‐1.14 MPa, respectively. Plants also higher stomatal safety margin (water difference between closure P ), needed a longer time reach dry‐down experiment, showed faster recovery gas exchange upon rehydration impaired signaling resulted canopy area conductance, which likely contributed delaying for plants hydraulic damage during drought. These findings clear association structural physiological changes improved against drought‐induced dysfunction mutant.

Language: Английский

Citations

1

Contrasting survival strategies for seedlings of two northern conifer species to extreme droughts and floods DOI

Katlyn A Schulz,

Alexandra Barry, Laura S. Kenefic

et al.

Tree Physiology, Journal Year: 2024, Volume and Issue: 44(10)

Published: Sept. 8, 2024

Lowland northern white-cedar (Thuja occidentalis L.) forests are increasingly exposed to extreme droughts and floods that cause tree mortality. However, it is not clear the extent which these events may differentially affect regeneration of cedar its common associate, balsam fir (Abies balsamea (L.) Mill.). To test this, we measured how seedlings were able avoid, resist recover from experimental drought flood treatments different lengths (8 66 days). Overall, found exhibited a strategy stress resistance growth recovery (resilience) moderate stress. Fir, on other hand, appears be adapted avoid overall lower resilience. In treatments, evidence stomatal behaviors. Cedar used available water quickly therefore experienced more than fir, but was survive at potentials > 3 MPa below key hydraulic thresholds. On employed conservative water-use avoided extremely low potential. response survival higher only reached 50% if 23.1 days flooding in contrast 7.4 reach mortality for fir. both floods, many stressed maintain partially brown canopies often survived stress, albeit with reduced growth, suggesting contrast, or had threshold-type responses they either full live little effect died reliance avoidance. Combined variable precipitation regimes, seasonal complex microtopography can provide safe sites forests, results inform conservation management lowland stands.

Language: Английский

Citations

1

Within‐leaf variation in embolism resistance is not a rule for compound‐leaved angiosperms DOI Creative Commons
Ian M. Rimer, Scott A. M. McAdam

American Journal of Botany, Journal Year: 2024, Volume and Issue: unknown

Published: Dec. 16, 2024

Abstract Premise Hydraulic segmentation, caused by the difference in embolism resistance across plant organs, provides a sacrificial layer of cheaper like leaves, to protect more costly such as stems, during drought. Within‐leaf hydraulic segmentation has been observed two compound‐leaved tree species, with leaflets being vulnerable than rachis or petiole. Many herbaceous species have compound and some that are associated pulvini at base lamina, which could provide an anatomical means preventing from spreading within leaf because higher number vessel endings pulvinus. Methods We used optical vulnerability method investigate whether differences were tissues six one deciduous leaves. Our selection included both palmately pinnately‐compound leaved each pulvinus leaflets. Results found considerable variation measured, but no evidence leaf. In pulvini, we major events crossing pulvinus, petiole into embolizing same water potential. Conclusions conclude within‐leaf resistance, is not universal phenomenon presence does barrier spread

Language: Английский

Citations

1

Weak link or strong foundation? Vulnerability of fine root networks and stems to xylem embolism DOI Creative Commons
Beatrice L. Harrison Day, Craig R. Brodersen, Timothy J. Brodribb

et al.

New Phytologist, Journal Year: 2024, Volume and Issue: unknown

Published: Sept. 12, 2024

Resolving the position of roots in whole-plant hierarchy drought-induced xylem embolism resistance is fundamental for predicting when species become isolated from soil water resources. Published research generally suggests that are most vulnerable organ plant vascular system, although estimates vary significantly. However, our knowledge root excludes fine (< 2 mm diameter) form bulk total absorptive surface area network and nutrient uptake. We measured stem vulnerability 10 major land clades (five angiosperms, three conifers, a fern lycophyte), using standardised situ methods (Optical Methods MicroCT). Mean across matched or exceeded stems all study species. In six these (one fern, one lycophyte, conifers angiosperm), were significantly more resistant than stems. No clear relationship was found between conduit diameter vulnerability. These results provide insight into hydraulic pathway at site uptake, challenge long-standing assumption

Language: Английский

Citations

0