A
well-preserved
left
theropod
metatarsal
I
from
the
Campanian
Mesaverde
Formation
of
Wyoming
is
described
and
identified
as
belonging
to
a
caenagnathid,
representing
first
occurrence
this
clade
formation.
The
specimen
unique
in
being
relatively
small,
but
featuring
suite
characters
(triangular
shaft,
less
constriction
between
shaft
distal
condyle,
spherical
condyle)
that
are
seen
larger
examples
Caenagnathidae
such
Anzu
wyliei
or
“Macrophalangia
canadensis”.
This
suggests
previously-observed
differences
small
large
caenagnathids
not
solely
result
allometry,
may
represent
phylogenetically
informative
variation.
Although
limited,
new
lends
some
support
hypothesis
canadensis”
does
Chirostenotes
pergracilis.
Furthermore,
important
establishing
presence
within
fauna,
which
remains
poorly
known.
Journal of Vertebrate Paleontology,
Journal Year:
2021,
Volume and Issue:
41(4)
Published: July 4, 2021
Tyrannosaurid
theropods
topped
the
terrestrial
food
chain
in
North
America
and
Asia
during
latest
Cretaceous.
Most
tyrannosaurids,
exemplified
by
Tyrannosaurus
rex,
had
deep
snouts,
thick
teeth,
large
jaw
muscles
that
could
generate
high
bite
forces.
They
coexisted
with
a
morphologically
divergent
group
of
long-snouted
relatives,
called
alioramins.
Qianzhousaurus
sinensis,
from
Maastrichtian
Ganzhou,
China,
is
largest
alioramin
yet
discovered,
but
has
only
been
briefly
described.
Here
we
present
detailed
osteological
description
holotype
cranium
mandible
Qianzhousaurus.
We
identify
several
new
autapomorphic
features
genus,
synapomorphies
unite
alioramins
(Qianzhousaurus,
Alioramus
altai,
remotus)
as
clade,
including
laterally
projecting
rugosity
on
jugal.
clarify
elongate
skull
involves
lengthening
anterior
palate
not
premaxilla,
reflected
posterior
bones
lower
jaw,
even
though
(orbit
lateral
temporal
fenestra)
are
proportionally
similar
to
deep-skulled
tyrannosaurids.
show
much
variation
among
species
consistent
growth
trends
other
A.
remotus,
represent
different
ontogenetic
stages
progressive
maturity,
across
which
signature
nasal
rugosites
became
less
prominent.
predict
represents
adult
level
maturity
for
alioramins,
propose
morphology
indicates
weaker
than
suggestive
differences
prey
choice
feeding
style.
PLoS ONE,
Journal Year:
2022,
Volume and Issue:
17(10), P. e0266648 - e0266648
Published: Oct. 19, 2022
Reconstructing
the
evolution,
diversity,
and
paleobiogeography
of
North
America’s
Late
Cretaceous
dinosaur
assemblages
require
spatiotemporally
contiguous
data;
however,
there
remains
a
spatial
temporal
disparity
in
data
on
continent.
The
rarity
vertebrate-bearing
sedimentary
deposits
representing
Turonian–Santonian
ecosystems,
relatively
sparse
record
dinosaurs
from
eastern
portion
continent,
present
persistent
challenges
for
studies
American
evolution.
Here
we
describe
an
assemblage
ornithomimosaurian
materials
Santonian
Eutaw
Formation
Mississippi.
Morphological
coupled
with
osteohistological
growth
markers
suggest
presence
two
taxa
different
body
sizes,
including
one
largest
ornithomimosaurians
known
worldwide.
regression
predicts
femoral
circumference
mass
individuals
similar
to
or
greater
than
that
large-bodied
ornithomimosaurs,
Beishanlong
grandis
,
Gallimimus
bullatus
.
paleoosteohistology
MMNS
VP-6332
demonstrates
individual
was
at
least
ten
years
age
(similar
B
[~375
kg,
13–14
old
death]).
Additional
pedal
elements
share
some
intriguing
features
yet
larger-body
size
closer
Deinocheirus
mirificus
ornithomimosaur
this
region
during
time
is
consistent
recent
discoveries
early-diverging,
ornithomimosaurs
mid-Cretaceous
strata
Laurasia
(
Arkansaurus
fridayi
).
smaller
taxon
represented
by
tibia
preserving
seven
cycles,
indicators
decreasing
growth,
belongs
approaching
somatic
maturity,
suggesting
co-existence
medium-
America.
provide
key
information
diversity
distribution
Appalachian
fit
broader
evidence
multiple
cohabiting
species
ecosystems
Laurasia.
The Anatomical Record,
Journal Year:
2022,
Volume and Issue:
306(7), P. 1918 - 1938
Published: Oct. 23, 2022
Abstract
Despite
the
long
history
of
research
in
late
Campanian
Judith
River
Formation
northern
Montana,
most
vertebrate
fossils
are
represented
by
fragmentary
remains,
making
precise
taxonomic
identifications
difficult.
Contrary
to
this,
partially
contemporaneous
Dinosaur
Park
Formation,
Alberta,
Canada
is
known
for
its
tremendous
fossil
preservation,
permitting
rigorous
studies
dinosaur
diversity,
evolution,
and
biostratigraphy.
Hadrosaurids
comprise
one
abundant
clades
but
affinities
hadrosaurid
specimens
remain
poorly
understood
Formation.
Corythosaurus
common
and,
date,
has
been
restricted
this
formation.
This
study
reports
first
definitive
from
which
were
discovered
on
two
private
ranches
Montana.
The
attribution
complete
skeleton
indicated
by:
wide
crest‐snout
angle,
presence
premaxilla‐nasal
fontanelle,
dorsoventrally
expanded
nasal,
laterally
exposed
ophthalmic
canal
laterosphenoid,
tall
neural
spines.
A
second
specimen
preserves
a
large
ilium
that
can
be
positively
identified
as
based
associated
skull,
now
hands.
recovered
Coal
Ridge
Member
approximately
time
equivalent
Thus,
discovery
extends
biogeographic
range
genus
establishes
framework
future
interformational
biostratigraphic
Late
Cretaceous
faunas
North
America.
Geological Society of America Bulletin,
Journal Year:
2023,
Volume and Issue:
136(7-8), P. 2689 - 2701
Published: Nov. 15, 2023
Abstract
The
Cretaceous
paleocommunities
of
North
America
preserve
a
rich
record
biodiversity
that
suggests
many
species
occupied
narrow
biogeographic
ranges
in
comparison
to
their
ecological
equivalents
extant
systems.
How
taxa
these
systems
partitioned
niches
and
structured
communities
can
be
difficult
determine
from
fossils
alone,
which
has
led
variety
hypotheses
concerning
diets
habitat
use.
Here,
we
examine
element
ratios
(Sr/Ca,
Ba/Ca)
the
enamel
suite
co-occurring
vertebrate
sampled
spatiotemporally
constrained
interval
Oldman
Formation
Alberta,
Canada,
reconstruct
trophic
structure,
use
δ13C,
δ18O,
87Sr/86Sr
compositions
test
for
niche
partitioning
among
hadrosaurids,
ceratopsids,
ankylosaurs.
We
also
previously
proposed
dietary
troodontid
theropods.
In
large
ornithischians,
find
Ba/Ca
Sr/Ca
are
consistent
with
herbivory,
hadrosaurs
distinct
ceratopsids
ankylosaurids
ranges,
pattern
is
indicative
differences
use/breadth,
plant
sources,
feeding
height.
mammals,
varanoid
lizards,
dromaeosaurids,
tyrannosaurids
gradient
lower
animal-dominant
omnivorous
faunivorous
diets.
Troodontids,
have
been
variably
hypothesized
as
either
faunivorous,
omnivorous,
or
herbivorous
due
unusual
dentition,
fall
between
those
ornithischians
dromaeosaurids.
From
multi-proxy
data,
interpret
troodontids
mixed-feeding
plant-dominant
omnivores.
These
proxies
represent
valuable
tool
understanding
community
ecology
ecosystems
hold
enormous
potential
future
research
paleobiology.
PeerJ,
Journal Year:
2021,
Volume and Issue:
9, P. e12640 - e12640
Published: Dec. 6, 2021
Borogovia
gracilicrus
is
a
small-bodied
theropod
dinosaur
from
the
Maastrichtian
(Upper
Cretaceous)
Nemegt
Formation
of
southern
Mongolia.
The
taxon
based
on
single
fragmentary
specimen
preserving
only
distal
part
hindlimbs.
morphology
shows
peculiar
combination
features,
some
which
are
traditionally
considered
troodontid
synapomorphies
and
others
unusual
for
Troodontidae
but
shared
with
other
maniraptoran
clades.
In
particular,
second
toe
B.
differs
troodontids
in
lacking
features
contribute
to
specialized
‘sickle-clawed’
toe,
here
termed
‘falciphoran
condition’,
dromaeosaurids
paravians,
such
as
strongly
compressed
falciform
ungual.
Phylogeny
reconstructions
intended
explore
affinities
consistently
support
its
referral
within
subclade
including
all
Late
Cretaceous
taxa.
placement
not
significantly
affected
by
combinations
hindlimb
or
homoplasy
elements
forming
falciphoran
condition.
supported
valid
distinct
troodontids,
Tochisaurus
Zanabazar
.
lack
ungual,
distinctive
interpreted
derived
specialization
among
retention
plesiomorphic
condition
non-paravian
theropods.
Acta Palaeontologica Romaniae,
Journal Year:
2024,
Volume and Issue:
20 (1), P. 77 - 86
Published: Feb. 11, 2024
Fragmentary
but
scientifically
important
pterosaur
fossils
have
been
reported
in
Lower
Cretaceous
strata
such
as
the
Jinju
and
Hasandong
formations
of
South
Korea.
Recently,
several
large
teeth
a
second
wing
phalanx
found
these
suggested
to
affinities
with
Boreopteridae,
clade
which
has
so
far
only
Yixian
Formation
China.
Here,
phylogenetic
proposed
boreopterid
specimens
are
reassessed,
using
two-dimensional
geometric
morphometric
shape
analyses.
The
analyses
Korean
different
morphospaces
from
Fm.
boreopterids,
plot
them
closer
anhanguerians.
An
isolated
does
close
Zhenyuanopterus
longirostris
morphospace
defined
by
first
two
principal
component
axes,
taxa
lineages
also
plotted
closely,
or
even
closer.
Thus,
there
seems
be
no
clear
evidence
that
belong
tooth
appear
very
those
established
members
this
clade.
Therefore,
hypothesis
some
can
referred
is
here
tentatively
rejected.
Research Square (Research Square),
Journal Year:
2024,
Volume and Issue:
unknown
Published: June 7, 2024
Abstract
A
new
troodontid
dinosaur,
Hypnovenator
sasayamaensis
gen.
et
sp.
nov.,
is
described
based
on
an
articulated
postcranial
skeleton
recovered
from
the
fluvial
deposits
of
Albian
Ohyamashimo
Formation
Sasayama
Group
in
Tambasasayama
City,
Hyogo
Prefecture,
Japan.
Hypnovenator
distinguished
other
troodontids
by
four
autapomorphies
and
a
combination
additional
features.
Our
phylogenetic
analysis
positions
as
oldest
one
most
basal
troodontines,
forming
monophyletic
clade
with
Gobivenator
mongoliensis.
The
discovery
suggests
that
small-bodied
maniraptorans
sleeping
posture
were
common
not
only
environments
volcanic
eolian
events
or
alluvial
systems
but
also
systems.
Geometric
morphometric
manual
unguals
shows
I
III
exhibit
considerable
morphological
variation
are
functionally
similar,
which
differs
those
non-troodontine
troodontids,
reflecting
transition
motion
within
Troodontinae.
has
mosaic
features
pes
related
to
cursoriality.
This
study
reveals
asymmetrical
arctometatarsus
occurred
Albian,
some
changes,
such
shorter
digit
IV
than
non-ungual
phalanges
digits
roller
joints
weakly
ginglymoid
articulation,
arose
during
early
Late
Cretaceous.
Canadian Journal of Earth Sciences,
Journal Year:
2024,
Volume and Issue:
61(11), P. 1127 - 1155
Published: Oct. 28, 2024
Late
Campanian
terrestrial
communities
of
western
Canada
are
best
known
from
the
fluvial–paralic
deposits
Dinosaur
Park
Formation
(DPF)
in
Provincial
(DPP),
Alberta.
However,
a
growing
list
localities
isolated
DPF
outcrops,
outside
DPP
area,
offers
glimpse
into
palaeocommunities
that
evolved
isochronously
with
biotas
greater
proximity
to
Western
Interior
Seaway.
Over
past
decade,
one
such
locality
was
explored
along
Lake
Diefenbaker
Saskatchewan
Landing
Park.
The
initial
palaeoecological
analysis
this
marginal
marine
community
based
on
palynomorph
and
vertebrate
microfossil
diversity
has
laid
foundation
for
current
study
its
monodominant
ceratopsian
bonebed.
latter
resulted
new
occurrences
Centrosaurus
apertus
elmisaurine
Citipes
elegans
incomplete
yet
diagnostic
specimens.
is
unequivocally
identified
by
parietal
bar
bearing
two
prominent
P1
P2
hooks,
which
expands
geographical
habitat
range
species
most
coastal
environment
DPF.
Furthermore,
presence
suggests
region
closer
age
lower
than
uppermost
DPP,
at
odds
previous
palynostratigraphic
interpretation.
faunal
composition
bonebed
also
supports
widely
distributed
metacommunity
across
these
deposits.
This
contribution
demonstrates
how
evidence
multiple
spatial
gradient,
beyond
temporal
gradient
available
within
alone,
picture
as
potential
model
system
biotic
turnover
response
sea
level
rise
geological
scale.
Journal of Vertebrate Paleontology,
Journal Year:
2021,
Volume and Issue:
41(5)
Published: Aug. 27, 2021
Despite
an
abundance
of
ornithomimid
fossils
from
the
Late
Cretaceous
Alberta,
Canada,
only
isolated
elements
are
known
upper
Maastrichtian
Scollard
Formation.
Ornithomimus
velox
and
Struthiomimus
sedens
two
taxa
recognized
penecontemporaneous
formations
in
U.S.A.,
but
incomplete
nature
these
specimens
lack
contemporary
material
Alberta
creates
a
gap
our
understanding
diversity
during
late
North
America.
Here,
I
describe
first
diagnostic
Formation
central
providing
new
information
about
taxonomic
composition
American
ornithomimids
Maastrichtian.
The
fossil,
articulated
forelimb,
exhibits
metacarpal
ratios
features
manual
unguals
that
support
its
referral
to
genus
Ornithomimus.
second
associated
hind
limb,
belongs
large-bodied
ornithomimid,
based
on
characteristics
pedal
is
assigned
Struthiomimus.
Based
designations,
stratigraphic
ranges
both
extended
Campanian
Dinosaur
Park
through
which
constitutes
more
than
10
million
years
time.
These
offer
knowledge
unusually
long
range
coupled
with
difficulty
diagnosing
either
specimen
species
underscores
need
for
review
taxonomy.
bioRxiv (Cold Spring Harbor Laboratory),
Journal Year:
2022,
Volume and Issue:
unknown
Published: March 25, 2022
Abstract
Reconstructing
the
evolution,
diversity,
and
paleobiogeography
of
North
America’s
Late
Cretaceous
dinosaur
assemblages
requires
spatiotemporally
contiguous
data;
however,
there
remains
a
spatial
temporal
disparity
in
data
on
continent.
The
rarity
vertebrate-bearing
sedimentary
deposits
representing
Turonian–Santonian
ecosystems,
relatively
sparse
record
dinosaurs
from
eastern
portion
continent,
present
persistent
challenges
for
studies
American
evolution.
Here
we
describe
an
assemblage
ornithomimosaurian
materials
Santonian
Eutaw
Formation
Mississippi.
Morphological
coupled
with
osteohistological
growth
markers
suggest
presence
two
taxa
different
body
sizes,
including
one
largest
ornithomimosaurians
known
worldwide.
regression
predicts
femoral
circumference
mass
individuals
similar
to
or
greater
than
that
large-bodied
ornithomimosaurs,
Beishanlong
grandis
Gallimimus
bullatus
.
paleohistology
MMNS
VP-6332
demonstrates
individual
was
at
least
11
years
age
(similar
B.
[∼375
kg,
13–14
old
death]).
Additional
pedal
elements
share
some
intriguing
features
ornithomimosaurs
yet
larger-body
size
closer
Deinocheirus
mirificus
ornithomimosaur
this
region
during
time
is
consistent
recent
discoveries
early-diverging,
mid-Cretaceous
strata
Laurasia
(
Arkansaurus
fridayi
).
smaller
taxon
represented
by
tibia
preserving
seven
cycles,
indicators
decreasing
growth,
belongs
near
reaching
somatic
maturity
larger
taxon,
suggesting
co-existence
medium-
America.
provide
key
information
diversity
distribution
Appalachian
fit
broader
evidence
multiple
cohabiting
species
ecosystems
Laurasia.